From ancient acorns to future forests, the story of how oaks evolved and the many ways they shape our world. An oak begins its life with the precarious journey of a pollen grain, then an acorn, then a seedling. A mature tree may shed millions of acorns, but only a handful will grow. One oak may then live 100 years, 250 years, or even 13,000 years. But the long life of an individual is only a part of these trees’ story. With naturalist and leading researcher Andrew L. Hipp as our guide, Oak Origins takes us through a sweeping evolutionary history, stretching back to a population of trees that lived more than 50 million years ago. We travel to the ancient tropical Earth to see the ancestors of the oaks evolving side by side with the dinosaurs. We journey from the oaks’ childhood in the once-warm forests of the Arctic to the montane cloud forests of Mexico and the broad-leaved evergreen forests of Southeast Asia. We dive into current research on oak genomes to see how scientists study genes’ movement between species and how oaks evolve over generations—spanning tens of millions of years. Finally, we learn how oak evolutionary history shapes the forests we know today, and how it may even shape the forests of the future. Oaks are familiar to almost everyone, and beloved. They are embedded in our mythology. They have fed us, housed us, provided wood for our ships and wine barrels and homes and halls, planked our roads, and kept us warm. Every oak also has the potential to feed thousands of birds, squirrels, and mice and host countless insects, mosses, fungi, and lichens. But as Oak Origins makes clear, the story of the oaks’ evolution is not just the story of one important tree. It is the story of the Tree of Life, connecting all organisms that have ever lived on Earth, from oaks’ last common ancestor to us.